Controls: mouse to place and drag. R or right-click rotates a piece while dragging. Click an empty tile to buy a turret; drag a turret onto one of the same level to merge. Arrow keys / WASD move around the upgrade tree, Enter buys.

Update 9/7 updated with some fixins. also added a fresh wipe in the options menu to delete old saves. added achievements, added time speed 1/2/3x in upper right. tried to fix some bugs that I noticed. 

Comments

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Pretty cool. The balancing is off as it's too easy to achieve high levels and then unlock all upgrades within 1-2 runs.

I'm surprised there aren't more comments here as this would have been quite well received just 2 years ago.